Within the last forty years, understanding of the early radiation of the gymnosperms has been greatly augmented by the description of numerous gymnospermous reproductive structures from the Late Devonian and Early Carboniferous. While a great deal of data have been amassed from anatomically preserved specimens from the Early Carboniferous, most specimens identified from Late Devonian sediments have been based on compression fossils. A suite of disarticulated gymnospermous specimens has been recovered from Late Devonian (latest Famennian) sediments from Hook Head, County Wexford, Ireland. Three different types of ovules, a pollen organ, and vegetative axes are anatomically preserved in pyrite and fusain. Two of the ovules have not been previously described and the third represents a new report of Hydrasperma Long. The two new ovules are radially symmetrical with four integumentary lobes. Both types of ovules display hydrasperman reproduction. The larger of the two ovules possesses an integument composed of three distinct layers, a feature not previously described for Devonian gymnosperms. The pollen organ is comprised of a cluster of four microsporangia, the organization of which is consistent with lyginopterid pteridosperms. Scanning electron micrographs of the vegetative axes show tracheid wall thickening patterns, which include scalariform-bordered pits, circular-bordered pits, and crowded pitting. These new ovules increase the total number of anatomically preserved ovules from the Devonian to five and the total number of ovules known from the Devonian to thirteen. The known diversity of gymnosperms in the latest Famennian suggests that future efforts in understanding the origin of this group should focus on Frasnian or even Givetian sediments.

Key words: anatomy, Devonian, gymnosperms, Ireland, Lyginopteridaceae, ovules