LIVSHULTZ, TATYANA. L.H. Bailey Hortorium, 462 Mann Library, Cornell University, Ithaca, NY 14853. - Systematics and evolution of ant-leaves in the genus Dischidia (Asclepiadaceae).
The genus Dischidia comprises approximately 80 species of
succulent epiphytic vines native to Southeast Asia. The genus is
divided into three sections based on leaf morphology: section
Dischidia with laminar leaves, section Conchophylla with
shell-shaped leaves, and section Ascidifera with
pitcher-leaves. Many species of all three sections frequently occur in
the nests of arboreal ants, and ants use the leaves of section
Ascidifera and section Conchophylla as nesting sites.
Both shell- and pitcher-leaves develop via differential growth of the
adaxial and abaxial leaf surfaces, leading to the hypothesis that
pitcher-leaves evolved from shell-leaves via increased curvature of
the leaf during development. I present combined cladistic analysis of
morphology and sequences of the second intron of the nuclear gene
Leafy. The results support the monophyly of Dischidia
s.l. and the sister-group relationship of Dischidia and
Hoya. The segregrate genera Dischidiopsis,
Oistonema, Conchophyllum, and Leptostemma should
be included in Dischidia s.l. while the monotypic genus
Micholitzia should be excluded. The analysis also shows that
Dischidia species which have both alternate and opposite
phyllotaxy form a clade, a grouping which has never been formally
proposed. Section Conchophylla plus section Ascidifera
form a monophyletic group, consistent with the evolution of
pitcher-leaves from shell-leaves.
Key words: ants, Asclepiadaceae, Dischidia, epiphytes, Leafy, systematics